By Gerhard Gottschalk (auth.)
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Coli were to need NADPH 2 and pentose phosphates in a ratio of 2: 1 additional reactions would not be required. The sum would, therefore, be: 3 glucose-6-phosphate + 6NADP~ 3 pentose-5-phosphate + 3C0 2 + 6NADPH 2 Now, consider a situation where there is a demand for more than 2NADPH 2 for each pentose phosphate made; the excess pentose phosphate would have to be removed. This is accomplished by the enzymes transketolase and transaldolase, which convert pentose phosphates back into hexose phosphates.
2. Enzyme reactions which allow the formation of ribose-5-phosphate and xylulose-5-phosphate from ribulose-5-phosphate. The enzymes involved are phosphoribose isomerase and ribulose-5-phosphate-3-epimerase. 3. Transaldolase and transketolase reactions, which catalyze the formation of hexose-6-phosphates from pentose-5-phosphates. 9 summarizes all of the reactions of the oxidative pentose phosphate cycle. First glucose-6-phosphate is oxidized to 6-phosphogluconate by a NADP-specific glucose-6-phosphate dehyqrogenase.
The amount of radioactive CO 2 formed per hour is plotted against the time. [CO H. Wang, I. Stern, C. M. Gilmour, S. Klungsoyr, D. J. Reed, J. J. Bialy, B. E. Christensen, and V. H. Cheldelin: J. Bacterial. ] Ribonucleotides and Deoxyribonucleotides 49 oxidative pentose phosphate pathway and 72% into the Embden-Meyerhof pathway. 10, which demonstrates the rate of release of radioactive carbon dioxide from cultures of E. coli growing with radioactive glucose labeled in various positions. As can be seen, carbon atoms 3 and 4 are the preferential sources of CO 2; in the degradation of glucose by the Embden-Meyerhof pathway C 3 and C4 of glucose are converted to carboxyl groups of pyruvate and released as CO 2 during acetyl-Co A formation.
